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    Thus, preQ0 is reduced to preQ1 and inserted in tRNAs by bTGT ( Fig. 1 A ). Two subsequent enzymatic steps, carried out by QueA and QueG, produce the final Q nucleoside (for recent review, see ref. 9 ). In Archaea, the tRNA guanosine (15) transglycosylase enzyme (aTGT; EC ) is homologous to the bTGT enzyme and exchanges the G at position 15 with preQ0 in nearly all tRNAs ( 15, 16 ). The preQ0 is then modified to G by different types of amidotransferases archaeosine synthase (ArcS QueF-like, and.

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